In the 28th century, Valerian and Laureline are special operatives charged with keeping order throughout the human territories. On assignment from the Minister of Defense, the two undertake a mission to Alpha, an ever-expanding metropolis where species from across the universe have converged over centuries to share knowledge, intelligence, and cultures.

Extreme athlete turned government operative Xander Cage comes out of self-imposed exile, thought to be long dead, and is set on a collision course with deadly alpha warrior Xiang and his team in a race to recover a sinister and seemingly unstoppable weapon known as Pandora's Box.

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